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This system is responsible for lifting the wings during the recovery stroke.
They are then oriented horizontally on the recovery stroke to slash through the water without pushing the animal back.
During movement, an individual cilium deforms using a high-friction power stroke followed by a low-friction recovery stroke.
The difference is that the recovery stroke is at 90 degrees to the power stroke, so that the cilia avoid hitting each other.
The detached myosin head then hydrolyses ATP, and performs a recovery stroke to restore its initial position.
Drag swimmers use a cyclic motion where they push water back in a power stroke, and return their limb forward in the return or recovery stroke.
In this discussion the brace is the one used for stability in stoppers or as a recovery stroke to avoid capsize, not the one used when breaking in or out.
In the case of flagella the motion is often planar and wave-like, whereas the motile cilia often perform a more complicated 3D motion with a power and recovery stroke.
Medusae swim by a form of jet propulsion: muscles, especially inside the rim of the bell, squeeze water out of the cavity inside the bell, and the springiness of the mesoglea powers the recovery stroke.
The vents are intended to allow for the passage of water during the recovery stroke, but prevent passage during power strokes due to the blade angle, attempting to lessen effort during recovery and improve kick efficiency.
Propulsion occurs on the back-stroke (when the hairs are erect) and far less energy is expended in moving the leg forward on the recovery stroke (when the hairs collapse against the leg and so reduce the resistance to movement).
The beat of each cilium has two phases: a fast "effective stroke," during which the cilium is relatively stiff, followed by a slow "recovery stroke," during which the cilium curls loosely to one side and sweeps forward in a counter-clockwise fashion.