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Roughly speaking, this represents the ratio of photosynthesising to respiring material in the plant.
The root cause of the temperature drop may have been the removal of CO from the atmosphere by photosynthesising bacteria and eukaryotes in the previous periods.
The reefs soon cease to grow as the poor light transmissions through the murky waters prevents the zooxanthellae actively photosynthesising underlining the vital life-sustaining role sunlight plays in this relationship.
But about 2 billion years ago, photosynthesizing microbes that lived on hydrogen began to break down water and expel oxygen as a byproduct.
Grazing livestock, usually not left to wander, would eat the grass and would minimize any grass growth while grass left alone would eventually grow to cover its own growing buds, preventing them from photosynthesizing and killing the plant.